Two tweakers likely to play again for the tourists but Glenn Maxwell's shoulder issue could keep him out of the final ODI
Aussie spin twins set for series finale
Australia look set to stick with two spinners for Sunday's fifth and final ODI against England in Manchester where the home side will be seeking an historic 5-0 whitewash.
Glenn Maxwell remains in doubt with the shoulder injury that ruled him out of the fourth match, with Kane Richardson a contender to come back into the side for his third match of the series.
Shaun Marsh, the 34-year-old veteran who is Australia's leading run-scorer in this series with two centuries, skipped the running drills but joined in slips practice and seems set to take his place at No.3.
Ashton Agar, one of the standout performers for the visitors on this tour, said the Australians would "have to bring our absolute best" to avoid a whitewash.
"It's going to be tough, but I absolutely believe we can beat them: we have no choice but to believe that we can beat them," said Agar.
"But we need to play our very best cricket against a side that's played really well four games in a row now. We have to bring our absolute best.
"We have plenty to gain out of (Sunday), plenty to gain, to just put things into practice and just leave it all out there.
"It's the last game (of the series), you go 100 per cent every game but I think it means more to us now to go as hard as we can and just leave nothing out there."
The spinners were Australia's best bowlers as England chased down 310 in the previous match in Durham, and Nathan Lyon said he felt the side "may have missed a trick" by not opting for two slow bowlers in the XI early.
Captain Tim Paine didn't use Lyon and Agar in tandem in Durham, rotating them from one end and the quicks at the other.
Lyon's 1-38 from seven overs complemented Agar's 2-48 from eight and helped produce Australia's most controlled passage of play with the ball.
"Personally, yeah I believe we potentially may have missed a trick (by not playing two spinners)," Lyon said in Manchester.
"But I'm not a selector, all I can do is keep putting my hand up to perform well and make them make a choice to pick me.
"When I get my opportunity on Sunday, if I do again, I've just got to make sure I keep going out there and bowling well and trying to improve, try to help this young side improve and understand the game."
Agar, who was also Australia's best with the ball when England unloaded for a world record score of 6-481, said he was personally proud of his response.
"I've experienced similar sorts of things to that and, yeah, it was difficult," Agar said.
" I came away with 1-70 (in 10 overs) and (Jonny) Bairstow absolutely smacked the one he got out on to midwicket.
"I was proud of the way I tried to fight back during that spell and tried to keep my cool. All you can do is stay calm and focus on the next ball.
"We've certainly learnt from that but we just have to keep moving forwards."
Lyon entered this tour knowing he was the second-choice spinner for the 50-over format behind Agar, and the 78-Test veteran with 306 wickets to his name was delighted to play his first ODI in two years.
"It's a great opportunity for me to get better, especially in white-ball cricket bowling at the likes of Stoinis, Finch, Maxwell in the nets.
"Some of those guys are the best batters in the world and if I'm bowling well against them in the nets then you only keep learning.
"And I can bounce ideas of Ash. I'm a big fan of Ashton Agar, so when we bowl in the nets together, we're always talking, we're enjoying bowling."
England's spinners have taken 19 wickets in four matches so far. Moeen Ali has eight and Adil Rashid, who went past Graeme Swann as England's most successful white-ball spinner, has 11.
The fifth and final ODI starts at 8pm AEST, with live coverage on 9Gem.
Qantas tours of the UK and Zimbabwe
Australia ODI squad: Tim Paine (c), Aaron Finch (vc), Ashton Agar, Alex Carey, Travis Head, Nathan Lyon, Glenn Maxwell, Shaun Marsh, Michael Neser, Jhye Richardson, Kane Richardson, D’Arcy Short, Billy Stanlake, Marcus Stoinis, Andrew Tye
England ODI squad: Eoin Morgan (c), Moeen Ali, Jonny Bairstow, Jake Ball, Sam Billings, Jos Buttler, Sam Curran, Tom Curran, Alex Hales, Craig Overton, Liam Plunkett, Adil Rashid, Joe Root, Jason Roy, David Willey, Mark Wood
Australia T20 squad: Aaron Finch (c), Alex Carey (vc), Ashton Agar, Travis Head, Nic Maddinson, Glenn Maxwell, Jhye Richardson, Kane Richardson, D’Arcy Short, Billy Stanlake, Marcus Stoinis, Mitchell Swepson, Andrew Tye, Jack Wildermuth
England T20 squad: Eoin Morgan (c), Moeen Ali, Jonny Bairstow, Jake Ball, Jos Buttler, Sam Curran, Tom Curran, Alex Hales, Chris Jordan, Liam Plunkett, Adil Rashid, Joe Root, Jason Roy, David Willey
Qantas Tour of the UK
June 7: Australia beat Sussex by 57 runs at Hove
June 9: Australia beat Middlesex by 101 runs at Lord's
June 13: England won by three wickets at The Oval
June 16: England won by 38 runs in Cardiff
June 19: England won by 242 runs at Trent Bridge
June 21: England won by six wickets in Durham
June 24: Fifth ODI, Old Trafford
June 27: Only T20, Edgbaston (D/N)
Qantas T20I tri-series Tour of Zimbabwe
July 1: Zimbabwe vs Pakistan
July 2: Pakistan vs Australia
July 3: Australia vs Zimbabwe
July 4: Zimbabwe vs Pakistan
July 5: Pakistan vs Australia
July 6: Australia vs Zimbabwe
July 8: Final